HOUSTON, Texas - The Southern Jaguars lost in heartbreaking fashion as Texas Southern freshman kicker Miguel Arana nailed a game-winning 46-yard field goal as time expired to give TSU the 24-21 win, handing the Jags their third conference loss of the season and their fifth loss overall.

The Jags struck first on a Christian Johnson pass to Damien Henderson to go ahead 7-0 in the first quarter, but the Tigers responded with a 14-point second quarter with a pair of rushing touchdowns. Jacob Washington scored the first to tie the game at seven apiece, and after a Southern three-and-out, Athean Renfro ran in a 15-yard touchdown to take the lead at 14-7.

Southern clawed their way back from a 14-point deficit in the second half. Running back Trey Holly ran in an 11-yard touchdown to tie the game at 21 with less than five minutes to go, but the TSU offense responded with a 10-play drive, draining the remainder of the clock and setting up Arana for the game-winning kick.

Jags head coach Marshall Faulk attempted to ice Arana with his final timeout, but Arana ended up nailing the kick anyway.

Despite the loss, Holly had a very big day on the ground. With the passing game limited to just 18 attempts on the afternoon, Holly racked up 35 total carries for 156 yards and a touchdown. Mike Franklin chipped in with seven carries and 44 yards.

Southern heads on the road again next week to take on Florida A&M in Tallahassee, Fla., in a matchup on Halloween.